董賽紅, 任國棟
(河北大學生命科學學院,河北省動物系統學與應用實驗室,河北保定071002)
中國云南爍甲屬分類研究及中國三新紀錄種(鞘翅目:擬步甲科:爍甲族)
董賽紅, 任國棟*
(河北大學生命科學學院,河北省動物系統學與應用實驗室,河北保定071002)
研究了中國云南擬步甲科Tenebrionidae爍甲族Amarygmini爍甲屬AmarygmusDalman,1823,發現中國3新紀錄種,越北爍甲A.tonkineusPic, 1922、絲角爍甲A.filicornis(Gravely, 1915)和腫腿爍甲A.nodicornis(Gravely, 1915),云南2新紀錄種,艾多爍甲A.adonis(Pic, 1922) 和中國爍甲A.sinensisPic, 1922,提供了該屬云南已知種檢索表、地理分布。檢視標本保存于河北大學博物館。
云南; 爍甲屬; 分類; 新紀錄
Dalman(1823)依據模式種ChrysomelamicansFabricius, 1794建立爍甲屬Amarygmus,該屬隸屬于鞘翅目Coleoptera擬步甲科Tenebrionidae爍甲族Amarygmini,目前全世界已知900余種(Bremer & Lillig,2014),是爍甲族種類最多的屬,其形態特征變化大,主要分布于東洋區、古北區東部、巴布亞、澳大利亞等太平洋島嶼地區。我國已知37種(包括本文新紀錄3種),分布于云南、上海、浙江、福建、江西、廣東、廣西、海南、四川、臺灣和香港。本文通過整理及鑒定采自中國云南的爍甲標本,發現中國3新紀錄種,云南2新紀錄種。編制了云南已知9種的分類檢索表。
研究標本保存于河北大學博物館。采用Bremer(2014)專業術語。
爍甲屬AmarygmusDalman,1823
AmarygmusDalman, 1823: 1823, 60 (Type species:ChrysomelamicansFabricius, 1794). Gebien 1920, 409-410; Bremer 2005b, 201.
DietysusPascoe, 1866: 1866, 486 (Type species:DietysusconfususPascoe, 1866). Synonymised by Gebien 1920, 410.
ElixotaPascoe, 1866: 1866, 475 (Type species:ElixotacupreaPascoe, 1866). Synonymised by Bremer 2001a, 65.
EuryperaPascoe, 1870: 1870, 106 (Type species:EuryperacupreaPascoe, 1870). Synonymised by Blackburn, 1893, 53.
AphyllocerusFairmaire, 1881: 1881a, 248; 1881b, 349 (Type species:AphyllocerusdecipiensFairmaire, 1881). Synonymised by Gebien 1920, 410.
AnacycusFairmaire, 1896: 1896, 33 (Type species:AnacycusnavicularisFairmaire, 1896). Synonymised to Elixota Pascoe, 1866 by Blair: 1929, 245.
PlatolenesGebien, 1913: 1913a, 420-421 (Type species:PlatolenesrufipesGebien, 1913). Synonymi-sed by Bremer 2001a, 69.
PseudamarygmusPic, 1915: 1915b, 9 (Type species:PseudamarygmustestaceipesPic, 1915). Synonymised by Bremer 2001a, 67.
ApelinaSaha, 1988: 1988, 429-430 (Type species:ApelinakeralaensisSaha, 1988). Synonymised by Bremer 2014, 3.
PlesiamarygmusMasumoto, 1989: 1989, 314 (Type species:DietysusovoideusFairmaire, 1882). Synonymised by Bremer 2005b, 205.
PyanirygmusPic, 1915: 1915b, 9 (Type species:PyanirygmuscorinthiusPic, 1915). Synonymised by Bremer 2005b, 206-208.
HyperamarygmusKaszab, 1964: 291-292 (Type species:HyperamarygmusantennalisKaszab, 1964). Synonymised by Bremer 2001a, 70.
OogetonMiwa, 1939: 412 (Type species:OogetonmakiiMiwa, 1939). Synonymised by Bremer 2005b, 209.
BecvaramarygmusMasumoto, 1999: 369 (Type species:DietysusnodicornisGravely, 1915). Synonymised by Bremer 2005b, 209.
PodamarygmusCarter, 1928: 287 (Type species:PodamarygmusalternansCarter, 1928). Synonymised by Bremer 2006, 37.
DryadigmusBremer, 2007: 26 (Type species:DryadigmuscechovskyiBremer, 2007). Synonymised by Bremer 2007, 26.
VarogetonBremer, 2014: 37 (Type species:DietysussubannulipesPic, 1922). Synonymised by Bremer 2014, 37.
屬征:身體卵圓形,有些種類延長,體色多樣;上顎端齒二裂狀,外緣具溝;觸角細長,6~10節略呈櫛齒狀;前足腿節近端部稍加粗,但缺齒及角狀突起。
分布:東洋區、古北區東部、巴布亞、澳大利亞等太平洋島嶼地區。

艾多爍甲Amarygmusadonis(Pic,1922),云南新紀錄
PlatolenesadonisPic, 1922: 1922a, 12; Kaszab 1980: 180.
Amarygmusadonis(Pic, 1922): Bremer 2001a, 57; Bremer, 2004: 11.
檢視標本:1ex.,云南景洪納板河保護區納板茶廠(森林,729 m),100.665 43°E,22.158 10°N,2009-V-16,孟令曾采。
分布:云南(景洪)、臺灣,越南,老撾,泰國,尼泊爾。
毛爍甲Amarygmuspilipes(Gebien,1913)
PlatolenespilipesGebien, 1913: 1913b, 42; Kaszab, 1980: 180.
AmarygmusformosanusPic, 1915: 1915a, 21. Synonymized by Ardoin, 1969: 126.
AmarygmuslongipillisPic, 1922: 1922b, 303. Synonymized by Bremer, 2001b: 84.
Amarygmuspilipes(Gebien, 1913): Bremer, 2001a: 57; Bremer, 2005a: 20; I. L?bl & A. Smetana, 2008: 216.
檢視標本:13ex.,云南滄源縣班洪鄉(1 130 m),2008-Ⅶ-16~18,徐吉山、郜振華采;1ex.,云南鎮康縣南傘鎮(1 000 m),2008-Ⅶ-19~21,徐吉山、周勇采;3ex.,云南景洪大渡崗,2014-Ⅳ-28,邱見玥采;1ex.,云南景谷,2006-Ⅷ-5~6,毛本勇等采;1ex.,云南金平阿得博,2006-Ⅶ-23~24,毛本勇等采;1ex.,云南耿馬縣城(1 100 m),2008-Ⅶ-15,徐吉山、郜振華采。
分布:云南(滄源、鎮康、景洪、景谷、金平、耿馬)、江西、福建、臺灣,越南,老撾,泰國,印度,緬甸,尼泊爾,馬來西亞。
粗點爍甲Amarygmuspunctatus(Pic,1922)
AnacycuspunctatusPic, 1922: 1922a: 11.
Elixotapunctate(Pic, 1922): Gebien, 1944: 513.
Amarygmuspunctatus(Pic, 1922): Bremer, 2001a: 57; Bremer, 2003: 72; I.L?bl & A. Smetana, 2008: 216.
檢視標本:3ex.,云南維西縣塔城鎮(2 012 m),2008-Ⅷ-7~9,徐吉山、郜振華采。
分布:云南(維西)、華南,老撾,泰國。
中國爍甲AmarygmussinensisPic,1922,云南新紀錄
AmarygmussinensisPic, 1922: 1922a: 11; Bremer, 2004: 33; I.L?bl & A. Smetana, 2008: 216.
檢視標本:1ex.,云南綠春平河鄉,2006-Ⅶ-26,毛本勇等采。
分布:云南(綠春)、浙江、福建。
光亮爍甲AmarygmusspeciosusDalman,1823
AmarygmusspeciosusDalman, 1823: 61; Bremer, 2003: 81; I.L?bl & A. Smetana, 2008: 216.
檢視標本:1ex.,N.Thailand, Doi Sang, 1990-Ⅴ-10~13,M.IT采。
分布:云南(德宏),老撾,泰國,印度,緬甸,尼泊爾。
越北爍甲AmarygmustonkineusPic,1922,中國新紀錄(圖版Ⅰ:1)
AmarygmuslacanusPic, 1924: 1924: 28; [syn.]: Bremer 2004: 39-40.
檢視標本:15ex.,云南金平阿得博,2006-Ⅶ-23~24,毛本勇等采。
分布:云南(金平),越南,老撾。
云南爍甲AmarygmusyunnanensisBremer,2011
AmarygmusyunnanensisBremer, 2011: 2011: 17.
檢視標本:未見。
分布:云南(西雙版納)。
絲角爍甲Amarygmusfilicornis(Gravely,1915),中國新紀錄(圖版Ⅰ:2)
DietysusfilicornisGravely, 1915: 532-553;ppl. XLIV, Fig. 14.
Amarygmusfilicornis(Gravely, 1915): Gebien 1944: 503.
DietysusatricolorPic, 1922: 1922b, 304.
Amarygmusatricolor(Pic, 1922): Gebien 1944: 502; [syn.]: Bremer 2005b: 217.
檢視標本:2ex.,云南金平十里村(草地,1 960 m),103.297 02°E,22.774 56°N, 2006-Ⅻ-16,白天,梁宏斌等采。
分布:云南(金平),越南,印度。
腫腿爍甲Amarygmusnodicornis(Gravely,1915),中國新紀錄(圖版Ⅰ:3~7)
DietysusnodicornisGravely, 1915: 533-534, pl. XLIV, Fig. 15.
Amarygmusnodicornis(Gravely, 1915): Gebien 1944: 504.
DietysusvitalisiPic, 1922: 1922b: 305.
Becvaramarygmusatricolor(Pic, 1922): Masumoto 1999: 369.
Amarygmusvitalisi(Pic, 1922): Gebien 1944: 505; [syn.]: Bremer 2005b: 214.
Amarygmus(Becvaramarygmus)nodicornis(Gravely, 1915); [stat. n.]: Bremer 2005b: 214.
檢視標本:2ex.,云南勐臘望天樹,2006-Ⅳ,朱笑愚采;1ex.,云南西雙版納勐侖,2014-Ⅹ,朱笑愚采。
分布:云南(西雙版納勐侖),老撾,印度。
該種主要鑒別特征是:雄性前足脛節膨大,且內側深凹,觸角2~6節端部明顯粗圓。日本學者Masumoto(1999)根據保存于法國自然歷史博物館的模式標本DietysusatricolorPic, 1922建立屬,僅記錄1種,德國學者Bremer(2005b)將其降為亞屬。

圖版Ⅰ 爍甲屬中國三新紀錄種Plate Ⅰ Three new record species of the genus Amarygmus Dalman, 1823 from China
1.越北爍甲AmarygmustonkineusPic, 1922; 2. 絲角爍甲Amarygmusfilicornis(Gravely, 1915); 3. 腫腿爍甲Amarygmusnodicornis(Gravely, 1915) ♂; 4. 腫腿爍甲Amarygmusnodicornis(Gravely, 1915) ♀; 5~7. 腫腿爍甲Amarygmusnodicornis(Gravely, 1915), 5.前足背面觀foreleg in dorsal view, 6. 前足腹面觀foreleg in ventral view, 7. 觸角antenna.
Ardoin P. 1969. Note synonymique (Col. Tenebrionidae)[J]. Bulletin de la Société entomologique de France, 74: 124-127.
Blair, KG. 1929. Some new species ofMyrmicophilousTenebrionidae (Col.)[J]. Zoologischer Anzeiger: 238-247.
Bremer HJ. 2001a. Revision der GattungAmarygmusDalman, 1823 und verwandter Gattungen. Ⅰ. Allgemeine Bemerkungen; Status einiger Gattungen affineAmarygmusDalman; neue Kombinationen von Arten der GattungAmarygmus(Coleoptera: Tenebrionidae: Alleculinae: Amagygmini)[J]. Coleoptera, 5: 57-80.
Bremer HJ. 2001b. Revision der GattungAmarygmusDalman, 1823 und verwandter Gattungen. Ⅱ. Neue Gattungen affineAmarygmusmit neuen Arten, sowie neue Arten und Synonyme[J]. Coleoptera, 5: 81-106.
Bremer HJ. 2003. Revision derGattungAmarygmusDalman, 1823 sowie verwandter Gattungen. ⅩⅥ. Erste Mitteilung über lang gestreckte Arten aus der orientalischen Region: Nachbeschreibungen und Abbildungen beschriebener Arten sowie Neubeschreibungen (Coleoptera: Tenebrionidae; Amarygmini)[J]. Annales Historico-Naturales Musei Nationalis Hungarici, 95: 37-105.
Bremer HJ. 2004. Revision der GattungAmarygmusDalman, 1823 sowie verwandter Gattungen. ⅩⅪ. Nachbeschreibungen, Neubeschreibungen und Illustrationen vonAmarygmus-Arten der orientalischen Region (Coleoptera; Tenebrionidae; Amarygmini)[J]. Acta Coleopterologica, 20(1): 7-86.
Bremer HJ. 2005a. Revision der GattungAmarygmusDalman, 1823 sowie verwandter Gattungen. ⅩⅩⅩⅢ. DieAmarygmus-Arten der orientalischen Region mit Makeln auf Flügeldecken. 2. Mitteilung (Coleoptera; Tenebrionidae; Amarygmini)[J]. Acta Coleopterologica, 21(2): 9-50.
Bremer HJ. 2005b. Revision der GattungAmarygmusDalman, 1823 sowie verwandter Gattungen. ⅩⅩⅩⅣ. Anmerkungen zu den GeneraAmarygmusDalman,BecvaramarygmusMasumoto,EumolpamarygmusPic,LobatopezusPic,OogetonKaszab undPyanirygmusPic (Insecta, Coleoptera, Tenebrionidae, Amarygmini, Chrysomelidae, Eumolpinae)[J]. Spixiana, 28(3): 199-221.
Bremer HJ. 2006. Revision der GattungAmarygmusDalman, 1823 und verwandter Gattungen. XLI. DieAmarygmus-Arten des SubgenusPodamarygmusCarter (Col.: Tenebrionidae: Amarygmini)[J]. Acta Coleopterologica, 22(1): 35-60.
Bremer HJ. 2007. Revision der GattungAmarygmusDalman, 1823 sowie verwandter Gattungen (Coleoptera: Tenebrionidae: Amarygmini). XLV. Neu- und Nachbeschreibungen vonAmarygmus-Arten der orientalischen Region[J]. Stuttgarter Beitr?ge zur Naturkunde, Serie A (Biologie), Nr. 707: 1-48.
Bremer HJ. 2011. Revision of the genusAmarygmusDalman and related genera. LIX. New elongateAmarygmusfrom Thailand, Laos, Vietnam and southern China (Coleoptera: Tenebrionidae: Amarygmini)[J]. Acta Coleopterologica, 27(2): 3-30.
Bremer HJ. 2014. Revision ofAmarygmusDalman, 1823 and related genera. part LXXII. The Amarygmini of Borneo (Coleoptera: Tenebrionidae), part Ⅳ. Species of the genusPlesiophthalmusand related genera (including species of Sumatra, Java and partially Peninsular Malaysia)[J]. Mitteilungen der Münchner Entomologischen Gesellschaft, 104: 31-83.
Bremer HJ, Lillig M. 2014. World Catalogue of Amarygmini, Rhysopaussini and Falsocossyphini (Coleoptera; Tenebrionidae)[J]. Mitteilungen der Münchner Entomologischen Gesellschaft, 104(Suppl.): 3-176.
Carter HJ. 1928. Revision of the Australian species of the generaCuris,NeocurisandTrachys, together with notes and descriptions of new species of other Coleoptera [J]. Proceedings of the Linnean Society of New South Wales, 53: 270-290.
Dalman JW. 1823. Analecta entomologica[M]. Holmiae: Typis Lindhianis: 104.
Fairmaire L. 1881a. Essai sur les coléoptères desles Viti (Fidgi) [J]. Annales de la Société entomologique de France, 1: 243-318.
Fairmaire L. 1881b. Diagnoses de coléoptères de la Mélanésie [J]. Le Naturaliste, 1: 348-349.
Fairmaire L. 1896. Hétéromères des l’Inde recueilles par M. Andrewes [J]. Annales de la Société entomologique de Belgique, 40: 6-60.
Gebien H. 1913a. Die Tenebrioniden der Philippinen [J]. The Philippine Journal of Science, Section D, 8: 373-433.
Gebien H. 1913b. H. Sauter’s Formosa-Ausbeute. Tenebrionidae (Coleopt.) [J]. Archiv für Naturgeschichte, 79A: 1-58.
Gebien H. 1920. Coleoptera, Tenebrionidae[M]// Nova Guinea, Résultats de l’expédition scientifique néerlandaise à la Nouvelle-Guinée en 1912 & 1913 sous les auspices de A. Franssen Herderschee. Vol. ⅩⅢ (3); Zoologie. Leiden: E. J. Brill: 213-500.
Gebien H. 1944. Katalog der Tenebrioniden[J]. Mitteilungen der Münchner Entomologischen Gesellschaft, 34: 497-555.
Gravely FH. 1915. Coleoptera, Ⅸ: Tenebrionidae[M]// Zoological results of the Abor Expedition, 1911-12. XLII. Coleoptera Ⅸ: Tenebrionidae. Records of the Indian Museum 8[1912-1922]: 519-536.
Kaszab Z. 1964. Tenebrioniden (Coleoptera) der Insel Amboina, gesammelt von A. M. R. Wegner[J]. Tijdschrift voor Entomologie, 107: 283-296.
Kaszab Z. 1980. Angaben zur Kenntnis der Tenebrioniden Nordvietnams (Coleoptera)[J]. Annales Historico-Naturales Musei Nationalis Hungarici, 72: 169-221.
Masumoto K. 1989.Plesiophthalmusand its allied genera (Coleoptera, Tenebrionidae, Amarygmini) (Part 4)[J]. The Japanese Journal of Entomology, 57: 295-317.
Masumoto K. 1999. Additions toPlesiophthalmusand its allied genera (Coleoptera, Tenebrionidae, Amarygmini) from East Asia, Part 1[J]. Elytra (Tokyo), 27(2): 353-370.
Miwa Y. 1939. Description on four new species belonging to the Family Tenebrionidae from Formosa[J]. Zoological Magazine (Tokyo), 5 (7): 412-415.
Pascoe FP. 1866. Notices of new or little-known genera and species of Coleoptera[J]. Journal of Entomology, 2: 443-493.
Pascoe FP. 1870. Additions to the Tenebrionidae of Australia[J]. The Annals and Magazine of Natural History including Zoology, Botany, and Geology, 5: 94-107.
Pic M. 1915a. Diagnoses d’Hétéromères[J]. Mélanges Exotico-entomologiques, 16: 14-24.
Pic M. 1915b. Genre nouveaux,espèces & variétés nouvelles[J]. Mélanges Exotico-entomologiques, 16: 2-13.
Pic M. 1922a. Nouveautés diverses[J]. Mélanges Exotico-entomologiques, 36: 1-32.
Pic M. 1922b. Sur les Hétéromères Amarygminae [Col.][J]. Bulletin de la Société zoologique de France, 47: 303-306.
Pic M. 1924. Nouveautés diverses[J]. Mélanges exotico-entomologiques, 41: 1-32.
Saha GN. 1988. A new genus of Amarygmini (Coleoptera: Tenebrionidae) from India[J]. Records of the Zoological Survey of India, 85 (3): 429-432.
TaxonomyoftheGenusAmarygmusDalman,1823fromYunnan,ChinaandThreeNewRecordSpeciesfromChina(Coleoptera:Tenebrionidae:Amarygmini)
DONG Saihong, REN Guodong*
(The Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University,Baoding, Hebei Province 071002, China)
The genusAmarygmusDalman, 1823 from Yunnan, China was studied in this paper. Three new record species from China,A.tonkineusPic, 1922,A.filicornis(Gravely, 1915), andA.nodicornis(Gravely, 1915), and two new record species from Yunnan province,A.adonis(Pic, 1922) andA.sinensisPic, 1922 were reported. The keys to species in Yunnan and geographical distribution were provided. Specimens are deposited in the Hebei University Museum.
Yunnan;Amarygmus; taxonomy; new record
10.11984/j.issn.1000-7083.20170208
2017-06-29接受日期2017-08-11
國家自然科學基金項目(31572309, 31402003); 河北省研究生創新資助項目(S2016004)
董賽紅(1977—), 博士, 講師, 從事昆蟲分類學研究, E-mail:dongsaihong@163.com
*通信作者Corresponding author, E-mail:gdren@hbu.edu.cn
Q959.229.9
A
1000-7083(2017)06-0697-05